| BONTHRON, Stuart Mcmillan ACTIVE | Director | 09 Oct 2025 | | England | May 1976 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| COLLINS, Peter John, Dr. ACTIVE | Director | 21 Nov 2024 | | Italy | July 1964 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| COOPER, Colin ACTIVE | Director | 19 Dec 2024 | | United Kingdom | June 1970 | Business & Innovation Directorate,, Rutherford Appleton Laboratory,, Science And Technologies Facilities Council (Stfc), Harwell Campus,, Didcot, OX11 0QX, United Kingdom |
| GREENE, Jonathan William ACTIVE | Director | 06 Jun 2024 | | United States | November 1970 | 4806, Holt Street, Bellaire, Texas 77401, United States |
| WEIDMANN, Damien, Dr ACTIVE | Director | 11 Jun 2015 | | England | August 1974 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| BELAL, Mohammed RESIGNED | Director | 11 Jun 2015 | 31 Dec 2019 | England | October 1987 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| BELL, Linda, Dr RESIGNED | Director | 28 Mar 2018 | 31 Oct 2021 | England | August 1957 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| BESTWICK, Timothy David, Dr RESIGNED | Director | 07 Jan 2015 | 15 Dec 2015 | England | March 1962 | 2.27, R71, Science And Technology Facilities Council, Harwell Oxford, Didcot, Oxfordshire, OX11 0QX, England |
| BLOXAM, Andrew RESIGNED | Director | 31 Jan 2019 | 20 Jul 2022 | England | October 1977 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| DENNY, David James Edmonds RESIGNED | Director | 15 Dec 2015 | 01 Dec 2022 | United Kingdom | December 1970 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| FLINT, Robert Charles RESIGNED | Director | 11 Apr 2023 | 09 Dec 2025 | United Kingdom | September 1966 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| FULLER, John Andrew RESIGNED | Director | 15 Dec 2015 | 20 Sep 2018 | United Kingdom | April 1961 | 2.27, R71, Science And Technology Facilities Council, Harwell Oxford, Didcot, Oxfordshire, OX11 0QX, England |
| HOLDEN, John Peter Stefano RESIGNED | Director | 05 Apr 2018 | 31 Jan 2019 | England | September 1966 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| KIRBY, Elizabeth Ann, Dr RESIGNED | Director | 15 Dec 2015 | 30 Aug 2023 | United Kingdom | April 1971 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| MUIR, Andrew Victor Graham, Dr RESIGNED | Director | 15 Dec 2015 | 20 Jul 2022 | England | October 1963 | Babraham Hall, Babraham, Cambridge, Cambridgeshire, CB22 3AT, United Kingdom |
| NILSSON, Hans Daniel RESIGNED | Director | 01 Jan 2020 | 01 Mar 2021 | United Kingdom | September 1955 | 29, Primrose Gardens, London, NW3 4UJ, England |
| RONAYNE, Kate RESIGNED | Director | 01 Dec 2022 | 01 May 2024 | England | April 1978 | 6 Zephyr Building, Eighth Street, Harwell Oxford, Didcot, OX11 0RL, England |
| SAGAIDACHNA, Anastasiia RESIGNED | Director | 20 Jul 2022 | 16 Oct 2023 | England | January 1990 | Foresight Group, 23 Floor The Shard, 32 London Bridge Street, London, SE1 9SG, England |