| ECCLESTON, Roger Soulsby, Professor ACTIVE | Director | 06 Feb 2025 | | United Kingdom | May 1965 | 4th Floor, 140 Aldersgate Street, London, EC1A 4HY, England |
| ELLIOT, Ruth Elizabeth ACTIVE | Director | 21 Oct 2024 | | United Kingdom | August 1977 | 4th Floor, 140 Aldersgate Street, London, EC1A 4HY, England |
| PECK, Simon Robert Weston ACTIVE | Director | 07 Jul 2026 | | England | April 1978 | 4th Floor, 140 Aldersgate Street, London, EC1A 4HY, England |
| VERNON, Paul Morgan, Mr. ACTIVE | Director | 14 Jan 2021 | | United Kingdom | September 1966 | 4th Floor, 140 Aldersgate Street, London, EC1A 4HY, England |
| BAINES, Linda Beatrice RESIGNED | Secretary | 06 Feb 2008 | 09 Feb 2010 | | | 5 The Maples, Grove, Wantage, Oxfordshire, OX12 0BA |
| PROSSER, Pauline RESIGNED | Secretary | 18 Dec 2007 | 06 Feb 2008 | | | 8 Upper Francis Street, Abertridwr, Caerphilly, Mid Glamorgan, CF83 4DU |
| ROBBINS, Gary Paul RESIGNED | Secretary | 09 Feb 2010 | 13 Oct 2015 | | | Rutherford Appleton Laboratory, Harwell Oxford, Didcot, Oxfordshire, OX11 0QX |
| AFFONSO, Mark Andrew RESIGNED | Director | 27 Jan 2015 | 30 Jul 2023 | England | April 1963 | Rutherford Appleton Laboratory, Harwell Oxford, Didcot, Oxfordshire, OX11 0QX, United Kingdom |
| BESTWICK, Timothy David RESIGNED | Director | 09 Feb 2010 | 07 Jul 2026 | England | March 1962 | Ukaea, Culham Science Centre, Abingdon, OX14 3DB, England |
| BROWN, Allison Louise RESIGNED | Director | 06 Aug 2024 | 21 Oct 2024 | England | March 1971 | 6th Floor, 125, London Wall, London, EC2Y 5AS, England |
| CHERRY, Robert Paul RESIGNED | Director | 18 Dec 2007 | 06 Feb 2008 | Wales | September 1967 | 82 Garrod Avenue, Dunvant, Swansea, West Glamorgan, SA2 7XQ |
| COX, Martin RESIGNED | Director | 01 Apr 2010 | 03 Jun 2013 | United Kingdom | December 1956 | Uk Atomic Energy Authority, K1 1 5 Culham Science Centre, Abingdon, Oxfordshire, OX14 3DB, United Kingdom |
| HOLLIS, Eric Thomas RESIGNED | Director | 01 Apr 2010 | 31 Jul 2014 | England | July 1949 | Uk Atomic Energy Authority, Culham Science Centre, Abingdon, Oxfordshire, OX14 3DB, United Kingdom |
| JACKSON, Andrew Philip RESIGNED | Director | 06 Feb 2008 | 01 Apr 2010 | England | May 1951 | 6 Sparvells, Eversley, Hook, Hampshire, RG27 0QG |
| JENKINSON, Antonia Scarlett RESIGNED | Director | 27 Feb 2019 | 18 Jan 2023 | England | July 1969 | Ukaea, Culham Science Centre, Abingdon, OX14 3DB, England |
| MOSS, Steven Yandell RESIGNED | Director | 31 Jul 2014 | 16 Dec 2014 | United Kingdom | April 1953 | Culham Science Centre, Abingdon, Oxfordshire, OX14 3DB, England |
| PARTRIDGE, Alan, Dr RESIGNED | Director | 30 Jul 2023 | 06 Feb 2025 | United Kingdom | May 1964 | 4th Floor, 140 Aldersgate Street, London, EC1A 4HY, England |
| PRIDHAM, Catherine Juliette RESIGNED | Director | 03 Jun 2013 | 27 Feb 2019 | United Kingdom | April 1966 | Culham Science Centre, Culham Science Centre, Abingdon, Oxfordshire, OX14 3DB, England |
| RONAYNE, Kate RESIGNED | Director | 18 Oct 2018 | 31 Oct 2019 | England | April 1978 | Rutherford Appleton Laboratory, Harwell Oxford, Didcot, OX11 0QX, England |
| STEWART, Gordon Bryant RESIGNED | Director | 06 Feb 2008 | 27 Jan 2015 | England | March 1966 | Rutherford Appleton Laboratory, Harwell Oxford, Didcot, Oxfordshire, OX11 0QX |
| TOWNS ANDREWS, Elizabeth, Dr RESIGNED | Director | 06 Feb 2008 | 09 Feb 2010 | United Kingdom | December 1958 | 24 Pool Lane, Lymm, Cheshire, WA13 9BJ |
| WALLACE, Iain RESIGNED | Director | 18 Jan 2023 | 22 Mar 2024 | Scotland | May 1981 | 6th Floor, 125, London Wall, London, EC2Y 5AS, England |
| WHITE, Stephen Henry RESIGNED | Director | 06 Feb 2008 | 01 Apr 2010 | United Kingdom | December 1950 | 8 Bridus Mead, Blewbury, Didcot, Oxfordshire, OX11 9PJ |