| BULLMORE, Edward Thomas, Professor ACTIVE | Director | 08 Dec 2016 | | United Kingdom | September 1960 | Herchel Smith Building For Brain & Mind Sciences, Cambridge Biomedical Campus, Cambridge, Cambs, CB2 0SZ, United Kingdom |
| JOHNSON, Anne Mandall, Professor Dame ACTIVE | Director | 03 Dec 2020 | | England | January 1954 | 41, Portland Place, London, W1B 1QH |
| MALIM, Michael Henry, Professor ACTIVE | Director | 27 Nov 2018 | | England | July 1963 | 2nd Floor, Borough Wing, Room Gh0402195, Guy's Hospital, London Bridge, Great Maze Pond, London, SE1 9RT, England |
| LACHMANN, Peter Julius, Professor RESIGNED | Secretary | 02 Mar 1998 | 10 Nov 2000 | | | 36 Conduit Head Road, Cambridge, Cambridgeshire, CB3 0EY |
| MANNING, Mary Elizabeth RESIGNED | Secretary | 10 Nov 2000 | 31 Dec 2009 | | | Flat 2 53 Millbank, London, SW1P 4RL |
| MUNN, Helen, Dr RESIGNED | Secretary | 31 Dec 2009 | 31 May 2019 | | | 41, Portland Place, London, W1B 1QH |
| ADAMS, David Harvey, Professor RESIGNED | Director | 18 Nov 2010 | 21 Dec 2013 | United Kingdom | February 1958 | Centre For Liver Research, 5th Floor Institute Of Biomedical Res, University Of Birmingham Medical School Edgbaston, Birmingham, B15 2TT |
| AGGLETON, John Patrick, Professor RESIGNED | Director | 16 Nov 2006 | 03 Dec 2009 | Wales | June 1955 | 36 Park Road, Whitchurch, Cardiff, South Glamorgan, CF14 7BR |
| ALBERTI, Kurt George Matthew Mayer RESIGNED | Director | 23 Nov 2000 | 21 Nov 2002 | | September 1937 | Flat 3 48 Percy Park, Tynemouth, North Shields, Tyne & Wear, NE30 4JX |
| ALMOND, Jeffrey William, Prof RESIGNED | Director | 07 Feb 2008 | 18 Nov 2010 | United Kingdom | June 1951 | 1541, Avenue Marcel Merieux, 69280 Marcy-L'Etoile Lyon, France, France, France |
| ASHMORE, Jonathan Felix, Professor RESIGNED | Director | 02 Dec 2014 | 05 Dec 2017 | Uk | April 1948 | University College London, Gower St, London, WC1E 6BT, Uk |
| AYNSLEY-GREEN, Albert RESIGNED | Director | 23 Nov 2000 | 20 Nov 2003 | | May 1943 | 54 Gibson Square, London, N1 0RA |
| BALKWILL, Frances Rosemary, Professor RESIGNED | Director | 16 Nov 2006 | 03 Dec 2009 | United Kingdom | March 1952 | 37 The Lexington, 40-56 City Road, London, EC1Y 2AN |
| BARNES, Peter John, Professor Sir RESIGNED | Director | 22 Nov 2007 | 18 Nov 2010 | England | October 1946 | Flat 2 28 Kempsford Gardens, London, SW5 9LH |
| BELL, John Irving, Professor Sir RESIGNED | Director | 20 Nov 2003 | 24 Nov 2011 | United Kingdom | July 1952 | Office Of Regius Prof, Richard Doll Bldg, Roosevelt Drive, Headington, Oxford, OX3 9DG |
| BESSER, Michael Gordon, Professor RESIGNED | Director | 23 Nov 2000 | 20 Nov 2003 | England | January 1936 | 61 Marlborough Place, London, NW8 0PT |
| BIGGS, Peter Martin, Professor RESIGNED | Director | 23 Nov 2000 | 20 Nov 2001 | United Kingdom | August 1926 | The Willows, London Road, St Ives, Cambridgeshire, PE27 5ES |
| BIRD, Colin Carmichael, Professor RESIGNED | Director | 21 Nov 2002 | 17 Nov 2005 | United Kingdom | March 1938 | 61 Lincolns Inn Fields, London, WC2A 3PX |
| BLACK, Carol Mary, Professor RESIGNED | Director | 21 Oct 2002 | 17 Nov 2005 | | December 1939 | Royal College Of Physicians, 11 St Andrews Place, London, NW1 4LE |
| BLOOM, Stephen Robert, Professor Sir RESIGNED | Director | 17 Nov 2004 | 22 Nov 2007 | England | October 1942 | 49 Hollycroft Avenue, Hampstead, London, NW3 7QJ |
| BOBROW, Martin, Professor RESIGNED | Director | 23 Nov 2000 | 21 Nov 2002 | | February 1938 | The Old School, High Street, Balsham, Cambridge, CB1 6DJ |
| BRADLEY, John Andrew, Professor RESIGNED | Director | 20 Nov 2003 | 16 Nov 2006 | United Kingdom | October 1950 | 4 Chaucer Close, Cambridge, Cambridgeshire, CB2 2TS |
| BRECKENRIDGE, Alasdair Muir, Sir RESIGNED | Director | 22 Nov 2012 | 03 Dec 2015 | United Kingdom | May 1937 | Cree Cottage, Feather Lane, Heswall, Wirral, CH60 4RL, Uk |
| BRODSKY, Frances Martha, Professor RESIGNED | Director | 27 Nov 2018 | 03 Dec 2019 | England | March 1955 | Division Of Biosciences, Med Sciences Bldg-1308, University College London, Gower Street, London, WC1E 6BT, England |
| BROPHY, Peter James, Professor RESIGNED | Director | 01 Jan 2009 | 03 Dec 2009 | Scotland | May 1949 | 31, Blacket Place, Edinburgh, Midlothian, EH9 1RJ |
| BUCHAN, Alastair Mitchell, Professor RESIGNED | Director | 18 Nov 2010 | 21 Dec 2013 | United Kingdom | October 1955 | John Radcliffe Hospital, 94 Headley Way, Headington, Oxford, Oxon, OX3 9DU |
| CALDICOTT, Fiona, Dame RESIGNED | Director | 23 Nov 2000 | 21 Nov 2002 | | January 1941 | Somerville College, Woodstock Road, Oxford, Oxfordshire, OX2 6HD |
| CANTRELL, Doreen Ann, Professor RESIGNED | Director | 20 Nov 2003 | 17 Nov 2004 | Scotland | January 1957 | 11 Osprey Avenue, Piperdam Fowlis, Dundee, DD2 5GB |
| CARTER, Yvonne Helen, Professor RESIGNED | Director | 16 Nov 2006 | 03 Dec 2009 | United Kingdom | April 1959 | Oaklands, 575a Westwood Heath Road, Coventry, West Midlands, CV4 8AA |
| CATTO, Graeme Robertson Dawson, Sir RESIGNED | Director | 23 Nov 2000 | 20 Nov 2001 | Scotland | April 1945 | 4 Woodend Avenue, Aberdeen, Aberdeenshire, AB15 6YL |
| CHILVERS, Edwin, Professor RESIGNED | Director | 22 Nov 2012 | 03 Dec 2015 | England | March 1959 | Department Of Medicine, Box 157 Addenbrookes Hospital, Hills Road, Cambridge, Cambs, CB2 0QQ, Uk |
| COHEN, Jonathan, Professor RESIGNED | Director | 21 Nov 2002 | 17 Nov 2005 | England | October 1949 | 27 Montpelier Crescent, Brighton, East Sussex, BN1 3JJ |
| COLLINS, Rory Edwards, Professor Sir RESIGNED | Director | 03 Dec 2015 | 27 Nov 2018 | England | January 1955 | Ndph Richard Doll Building, Old Road, Headington, Oxford, Oxon, OX3 7LF, United Kingdom |
| CONNELL, John Muir Cochrane, Professor RESIGNED | Director | 22 Nov 2007 | 18 Nov 2010 | Scotland | October 1954 | 68, Newlands Road, Glasgow, G43 2JH |
| CORNER, Jessica Lois, Professor Dame RESIGNED | Director | 05 Dec 2017 | 03 Dec 2019 | United Kingdom | March 1961 | 41, Academy Of Medical Sciences, Portland Place, London, W1B 1QH, England |
| CRAFT, Alan William, Professor Sir RESIGNED | Director | 17 Nov 2004 | 22 Nov 2007 | United Kingdom | July 1946 | 2 Ruthven Court, 10 Adderstone Crescent, Newcastle Upon Tyne, Tyne &Wear, NE2 2HH |
| CRITCHLEY, Hilary Octavia Dawn, Professor RESIGNED | Director | 05 Dec 2017 | 03 Dec 2019 | Scotland | March 1957 | Centre For Reproductive Health, University Of Edinburgh, Queens Medical Research Institute, Edinburgh, EH16 4TJ, Scotland |
| DAY, Chris Paul, Professor RESIGNED | Director | 02 Dec 2014 | 05 Dec 2017 | England | February 1960 | 11, Percy Park, North Shields, Tyne And Wear, NE30 4LA |
| DAY, Christopher Paul, Prof RESIGNED | Director | 20 Nov 2008 | 24 Nov 2011 | United Kingdom | February 1960 | The Medical School, Framlington Place, Newcastle Upon Tyne, Uk, NE2 4HH, United Kingdom |
| DELL, Anne, Professor RESIGNED | Director | 02 Dec 2014 | 27 Nov 2018 | United Kingdom | September 1950 | Imperial College London, South Kensington, London, United Kingdom |
| DELPY, David Thomas, Professor RESIGNED | Director | 17 Nov 2004 | 31 Aug 2007 | England | August 1948 | 9 Keswick Avenue, Merton Park, London, SW19 3JE |
| DEZATEUX, Carol Anne, Professor RESIGNED | Director | 24 Nov 2011 | 02 Dec 2014 | United Kingdom | May 1954 | 30, Guilford Street, London, WC1N 1EH, United Kingdom |
| DOLLERY, Colin Terence, Professor Sir RESIGNED | Director | 20 Nov 2001 | 17 Nov 2005 | United Kingdom | March 1931 | 101 Corringham Road, London, NW11 7DL |
| DOLPHIN, Annette Catherine, Professor RESIGNED | Director | 03 Dec 2015 | 27 Nov 2018 | England | April 1951 | University College London, Gower St, London, WC1E 6BT, United Kingdom |
| DOMINICZAK, Anna Felicja, Professor Dame RESIGNED | Director | 27 Nov 2018 | 03 Dec 2019 | Scotland | August 1954 | Wolfson Medical School Building, University Of Glasgow, University Avenue, Glasgow, G12 8QQ, Scotland |
| DOMINICZAK, Anna Felicja, Professor Dame RESIGNED | Director | 16 Feb 2012 | 02 Dec 2014 | Scotland | August 1954 | Wolfson Medical School Building, University Of Glasgow, University Avenue, Glasgow, Strathclyde, G12 8TA, Scotland |
| DUNNETT, Stephen Bruce, Professor RESIGNED | Director | 21 Nov 2013 | 08 Dec 2016 | Uk | January 1950 | Cardiff University, School Of Biosciences, Museum Avenue, Cardiff, South Glamorgan, CF10 3AX |
| EDWARDS, Christopher, Professor RESIGNED | Director | 02 Mar 1998 | 20 Nov 2001 | | February 1942 | 6 Kensington Terrace, Newcastle Upon Tyne, Tyne & Wear, NE1 7RU |
| FAIRLAMB, Alan Hutchinson, Professor RESIGNED | Director | 18 Nov 2010 | 21 Nov 2013 | United Kingdom | April 1947 | Wellcome Trust Biocentre, Dow Street, University Of Dundee, Dundee, Angus, DD1 5EH |
| FELDMANN, Marc, Sir RESIGNED | Director | 22 Nov 2007 | 18 Nov 2010 | England | December 1944 | 5, Durham Terrace, London, W2 5PB |
| FISHER, Elizabeth Mary Claire, Professor RESIGNED | Director | 08 Dec 2016 | 03 Dec 2019 | United Kingdom | September 1959 | 41, Portland Place, London, W1B 1QH |
| FITZGERALD, Maria, Professor RESIGNED | Director | 22 Nov 2012 | 03 Dec 2015 | United Kingdom | April 1953 | University College London, Gower Street, London, WC1E 6BT, United Kingdom |
| FRACKOWIAK, Richard Stanislaus Joseph RESIGNED | Director | 23 Nov 2000 | 20 Nov 2003 | | March 1950 | 32 Airedale Avenue, London, W4 2NW |
| FRANKLYN, Jayne Agneta, Professor RESIGNED | Director | 22 Nov 2012 | 02 Dec 2014 | Great Britain | July 1956 | 41, Portland Place, London, W1B 1QH, England |
| FURR, Barrington John Albert, Professor RESIGNED | Director | 17 Nov 2004 | 22 Nov 2007 | | November 1943 | Cancer Research Uk, 61 Lincoln's Inn Fields, London, WC2A 3PX |
| GOLDBERG, David Paul Brandes, Sir RESIGNED | Director | 23 Nov 2000 | 20 Nov 2003 | | January 1934 | 7 Woodhall Drive, Dulwich, London, SE21 7HJ |
| GRIFFIN, George Edward, Professor RESIGNED | Director | 01 Feb 2014 | 27 Jun 2018 | United Kingdom | February 1947 | St Georges University Of London Infection And Immu, Cranmer Terrace, Tooting, London, SW17 0RE, United Kingdom |
| GRIFFIN, George Edward, Professor RESIGNED | Director | 20 Nov 2003 | 16 Nov 2006 | United Kingdom | February 1947 | 8 Buxton Drive, New Malden, Surrey, KT3 3UZ |
| HART, Graham John, Professor RESIGNED | Director | 21 Nov 2013 | 08 Dec 2016 | United Kingdom | June 1957 | 61, Lansdowne Drive, London, Greater London, E8 3EP, United Kingdom |
| HASLETT, Christopher, Prof RESIGNED | Director | 17 Nov 2004 | 22 Nov 2007 | | April 1953 | 12 Chambers Crescent, Edinburgh, EH9 1TS |
| HIGGINS, Christopher Francis, Professor RESIGNED | Director | 23 Nov 2000 | 20 Nov 2003 | | June 1955 | Pipal Cottage, Water Eaton, Oxford, Oxfordshire, OX2 8HE |
| HILL, Raymond George, Professor RESIGNED | Director | 10 Feb 2011 | 21 Nov 2013 | Uk | December 1945 | 41, Portland Place, London, W1B 1QH |
| HOLGATE, Stephen Townley, Professor RESIGNED | Director | 01 Jan 2009 | 24 Nov 2011 | England | May 1947 | Level B, South Academic Block, South Block, Southampton General Hospital, Southampton, Hampshire, SO16 6YD, England |
| HORTON, Richard Charles, Dr RESIGNED | Director | 03 Dec 2009 | 22 Nov 2012 | Uk | December 1961 | 32, Jamestown Road, London, NW1 7BY |
| HUMPHRIES, Martin James, Professor RESIGNED | Director | 22 Nov 2012 | 05 Dec 2017 | United Kingdom | November 1958 | Michael Smith Building, Oxford Road, Manchester, M13 9PT, Uk |
| HUMPHRIES, Martin James, Professor RESIGNED | Director | 17 Nov 2005 | 20 Nov 2008 | United Kingdom | November 1958 | 30 Thorn Road, Bramhall, Stockport, Cheshire, SK7 1HQ |
| IREDALE, John Peter, Professor RESIGNED | Director | 24 Nov 2011 | 02 Dec 2014 | Scotland | November 1960 | 47, Little France Crescent, Edinburgh, Mid Lothian, EH16 4TJ, Uk |
| IVERSEN, Susan Diana, Professor RESIGNED | Director | 18 Nov 2010 | 02 Dec 2014 | Uk | February 1940 | 41, Portland Place, London, W1B 1QH |
| IVERSEN, Susan Diana, Professor RESIGNED | Director | 17 Nov 2004 | 22 Nov 2007 | United Kingdom | February 1940 | University Of Oxford, Wellington Square, Oxford, Oxfordshire, OX1 2JD |
| JACOBS, Patricia Ann, Professor RESIGNED | Director | 23 Nov 2000 | 20 Nov 2001 | | October 1934 | 55 Old Road, Alderbury, Salisbury, Wiltshire, SP5 3AR |
| JOHNSON, Anne Mandall, Professor Dame RESIGNED | Director | 27 Jun 2018 | 03 Dec 2019 | England | January 1954 | Ucl Institute For Global Health, The Mortimer Market Centre, Mortimer Market (Off Capper Street), London, WC1E 6JB, England |
| JOHNSTONE, Eve Cordelia, Professor RESIGNED | Director | 20 Nov 2003 | 16 Nov 2006 | United Kingdom | September 1944 | 9 Hallhead Road, Edinburgh, Lothian, EH16 5QJ |
| JONES, Roger Hugh, Professor RESIGNED | Director | 08 Dec 2009 | 09 Jan 2012 | United Kingdom | November 1948 | Kcl Dept Of General Practice, Capital House, Weston Street, London, SE1 |
| JONES, Roger Hugh, Professor RESIGNED | Director | 21 Nov 2002 | 17 Nov 2005 | United Kingdom | November 1948 | 56 Scotts Sufferance Wharf, Mill Street, London, SE1 2DE |
| KAPUR, Shitij, Professor RESIGNED | Director | 26 Apr 2012 | 02 Dec 2014 | United Kingdom | September 1964 | 16, De Crespigny Park, Denmark Hill, London, SE5 8AF, England |
| KARET, Fiona Eve, Professor RESIGNED | Director | 22 Nov 2012 | 03 Dec 2015 | England | March 1961 | Cambridge Institute For Medical Research, Addenbrookes Hospital, Hills Road, Cambridge, Cambs, CB2 0QQ, England |
| KENDELL, Robert Evan, Dr RESIGNED | Director | 02 Mar 1998 | 23 Nov 2000 | | March 1935 | 3 West Castle Road, Edinburgh, EH10 5AT |
| KENNARD, Christopher, Professor RESIGNED | Director | 17 Nov 2004 | 22 Nov 2007 | United Kingdom | January 1946 | 1 Pembroke Square, London, W8 6PA |
| KHAW, Kay-Tee Tee, Professor RESIGNED | Director | 01 Jan 2010 | 22 Nov 2012 | United Kingdom | October 1950 | Clinical Gerontology Unit Box 251, Addensbrooke's Hospital, Canbridge, Cambridgeshire, CB22 2QQ |
| KINMONTH, Ann-Louise, Professor RESIGNED | Director | 23 Nov 2000 | 21 Nov 2002 | | January 1951 | 1 Cambridge Road, Great Shelford, Cambridgeshire, CB2 5JE |
| KIRKWOOD, Thomas Burton Loram, Professor RESIGNED | Director | 21 Nov 2002 | 17 Nov 2005 | United Kingdom | July 1951 | Roughlees, Ewesley, Morpeth, Northumberland, NE61 4PH |
| KRZYSZTOF BORYSIEWICZ, Sir Leszek, Professor RESIGNED | Director | 20 Nov 2001 | 17 Nov 2004 | | April 1951 | 90 Grange Road, Ealing, London, W5 3PJ |
| LACHMANN, Peter Julius, Professor RESIGNED | Director | 02 Mar 1998 | 21 Nov 2002 | England | December 1931 | 36 Conduit Head Road, Cambridge, Cambridgeshire, CB3 0EY |
| LANYON, Lance Edward, Professor RESIGNED | Director | 21 Nov 2002 | 17 Nov 2004 | | January 1944 | 9 School Lane, Bricket Wood, St. Albans, Hertfordshire, AL2 3XS |
| LASKEY, Ronald Alfred, Professor RESIGNED | Director | 22 Nov 2007 | 22 Nov 2012 | England | January 1945 | Department Of Zoology, Downing Street, Cambridge, CB2 3EJ, United Kingdom |
| LAUDER, Ian, Professor RESIGNED | Director | 17 Nov 2005 | 18 Nov 2010 | England | June 1946 | Paddock House, Illston On The Hill, Leicester, Leicestershire, LE7 9EG |
| LAWLOR, Deborah Anne, Professor RESIGNED | Director | 16 Dec 2014 | 08 Dec 2016 | Uk | April 1963 | Mrc Ieu At The University Of Bristol, Oakfield House, Oakfield Grove, Bristol, BS8 2BN, Uk |
| LECHLER, Robert Ian, Professor Sir RESIGNED | Director | 03 Dec 2015 | 03 Dec 2020 | United Kingdom | December 1951 | Kings Health Partners, Ground Floor Counting House Guys Hospital, St Thomas' Street, London, SE1 9RT, United Kingdom |
| LECHLER, Robert Ian, Professor RESIGNED | Director | 18 Nov 2010 | 21 Nov 2013 | United Kingdom | December 1951 | Room 1.4 Hodgkin Bldg, Guy's Hospital, New Cohen Street, London, SE1 1UL, Uk |
| LECHLER, Robert Ian, Professor RESIGNED | Director | 17 Nov 2005 | 20 Nov 2008 | United Kingdom | December 1951 | 78 Woodstock Road, Chiswick, London, W4 1EQ |
| LEE, Melanie Georgina, Dr RESIGNED | Director | 16 Dec 2013 | 08 Dec 2016 | Uk | July 1958 | Wellcome Gibbs Building, 215 Euston Road, London, NW1 2BE |
| LEES, Andrew John, Professor RESIGNED | Director | 22 Nov 2012 | 03 Dec 2015 | England | September 1947 | 1, Wakefield Street, London, WC1N 1PJ, England |
| LEIGH, Irene May, Prof RESIGNED | Director | 01 Jan 2009 | 24 Nov 2011 | United Kingdom | April 1947 | 6, Balruddery Meadows, Dundee, Angus, DD2 5LJ |
| LEMOINE, Nicholas Robert, Professor RESIGNED | Director | 03 Dec 2009 | 22 Nov 2012 | England | December 1957 | The Anatomy Building, Barts And The London Scholl Of Medicine, Charterhouse Square, London, EC1M 6BQ |
| LEMON, Roger Nicholas, Professor RESIGNED | Director | 24 Nov 2011 | 02 Dec 2014 | Uk | March 1946 | Sobell Department, Ucl Institute Of Neurology, Queen Square, London, WC1N 3BG, Uk |
| LOMAS, David Arthur, Professor RESIGNED | Director | 27 Nov 2018 | 03 Dec 2019 | England | February 1962 | University College London, 1st Floor, Maple House, 149 Tottenham Court Road, London, W1T 7JA, England |
| LOVELL BADGE, Robin Howard, Dr RESIGNED | Director | 22 Nov 2007 | 18 Nov 2010 | United Kingdom | June 1953 | 74, Ledbury Road, London, W11 2AH |
| MACSWEEN, Roderick Norman Mciver, Professor Sir RESIGNED | Director | 23 Nov 2000 | 21 Nov 2002 | | February 1935 | 32 Calderwood Road, Newlands, Glasgow, G43 2RU |
| MANNING, Mary Elizabeth RESIGNED | Director | 10 Nov 2000 | 20 Nov 2000 | | November 1947 | Flat 36, 1 Ambrosden Avenue, London, SW1P 1QQ |
| MANT, David, Professor RESIGNED | Director | 07 Jan 2015 | 03 Dec 2015 | Uk | November 1949 | Department Of Primary Care, Radcliffe Observatory Quarter, Oxford, Oxfordshire, OX2 7EY, Uk |