| WARREN, Louise Ann ACTIVE | Secretary | 01 Feb 2015 | | | | Harpenden, Herts, AL5 2JQ |
| GRAY, Julie Elizabeth, Professor ACTIVE | Director | 10 Feb 2022 | | England | July 1962 | Harpenden, Herts, AL5 2JQ |
| HOLTHUIS, Johannes Vincentius Maria ACTIVE | Director | 11 May 2021 | | Netherlands | July 1962 | Harpenden, Herts, AL5 2JQ |
| KENDALL, Peter Ashley, Sir ACTIVE | Director | 10 Dec 2024 | | United Kingdom | May 1960 | Harpenden, Herts, AL5 2JQ |
| LELLIOTT, John George ACTIVE | Director | 03 Aug 2026 | | England | August 1952 | Harpenden, Herts, AL5 2JQ |
| LOBLEY, Matthew, Professor ACTIVE | Director | 04 Jan 2024 | | United Kingdom | October 1965 | Harpenden, Herts, AL5 2JQ |
| O'LEARY, Paul Simon ACTIVE | Director | 09 Jul 2026 | | United Kingdom | August 1967 | Harpenden, Herts, AL5 2JQ |
| BARNES, William RESIGNED | Secretary | | 31 Aug 1996 | | | Flat 4 The Manor House, Rothamsted Estate, Harpenden, Herts, AL5 2BE |
| JAMES, Stephen RESIGNED | Secretary | 19 Nov 2003 | 01 Feb 2015 | | | 7 Chaseside Close, Cheddington, Bedfordshire, LU7 0SA |
| THOMAS, Peter Stuart RESIGNED | Secretary | 01 Sep 1996 | 19 Nov 2003 | | | Richmond Lodge, Rhinefield Road, Brockenhurst, Hampshire, SO42 7SQ |
| BAILEY, Mark Jonathan, Professor RESIGNED | Director | 20 Jul 2007 | 12 May 2015 | England | March 1957 | Brookvale House 19 Faringdon Road, Stanford In The Vale, Oxfordshire, SN7 8NN |
| BALMFORD, Andrew Paul, Professor RESIGNED | Director | 08 Sep 2021 | 10 Dec 2025 | United Kingdom | March 1963 | Harpenden, Herts, AL5 2JQ |
| BARDGETT, Richard David, Professor RESIGNED | Director | 18 Nov 2008 | 05 Feb 2019 | England | November 1964 | 139, Main Street, Warton, Carnforth, Lancashire, LA5 9QF |
| BAULCOMBE, David Charles, Sir RESIGNED | Director | 10 Dec 2013 | 10 Feb 2022 | United Kingdom | April 1952 | Department Of Plant Sciences, Downing Street, Cambridge, CB2 2EA, United Kingdom |
| BEDDINGTON, John Rex, Sir RESIGNED | Director | 13 May 2014 | 10 Dec 2024 | United Kingdom | October 1945 | Harpenden, Herts, AL5 2JQ |
| BIRCH, Graham John, Dr RESIGNED | Director | 01 Jul 2007 | 12 May 2015 | England | June 1960 | Orchard House Orchard House Lane, Off Holywell Lane, St. Albans, Hertfordshire, AL1 1BX |
| BRIGHTMAN, David Kenneth, Dr RESIGNED | Director | 22 Feb 2006 | 24 Sep 2013 | England | August 1954 | Gaydon Hill Farm, Gaydon, Warwick, Warwickshire, CV35 0HQ |
| BROOKS, Russell Leon RESIGNED | Director | 10 Feb 2015 | 04 Dec 2019 | England | May 1972 | Harpenden, Herts, AL5 2JQ |
| BROWN, Valerie Kathleen, Dr RESIGNED | Director | 26 Jun 2002 | 31 Jul 2004 | | May 1944 | Le Tour,9 Meadow Rise, Lacey Green, Princes Risborough, Buckinghamshire, HP27 0QY |
| CALVERT, Michael RESIGNED | Director | 30 Jun 2000 | 20 Mar 2003 | | May 1950 | Iscennen House, 8 Chapel Lane Gaddesby, Leicester, Leicestershire, LE7 4WB |
| CHAMBERLAIN, Philip Walter RESIGNED | Director | 30 Jun 2000 | 31 Dec 2012 | England | September 1951 | Crowmarsh Battle Farm, 84 Preston Crowmarsh, Wallingford, Oxon, OX10 6SL |
| COCKING, Edward Charles Daniel, Professor RESIGNED | Director | | 01 May 2013 | United Kingdom | September 1931 | Life Science Department, University Of Nottingham, University Park, Nottinghamshire, NG7 2RD |
| DOUBLEDAY, Oliver Peter, Dr RESIGNED | Director | 07 Dec 2015 | 10 Dec 2024 | England | February 1952 | Harpenden, Herts, AL5 2JQ |
| DOYLE, Peter, Doctor RESIGNED | Director | 30 Sep 1992 | 30 Apr 1998 | | September 1938 | Twin Oaks Crawley Drive, Camberley, Surrey, GU15 2AA |
| DRUMMOND, Caroline Jane RESIGNED | Director | 01 Jul 2007 | 12 May 2015 | England | June 1963 | Butterwell, Horningtops, Liskeard, Cornwall, PL14 3QD |
| ELLIS, Richard Marriott RESIGNED | Director | 23 Feb 2005 | 24 Nov 2008 | England | January 1955 | The Grange, Brinton, Melton Constable, Norfolk, NR24 2QH |
| ELVES, Michael William, Dr RESIGNED | Director | 02 Jul 1999 | 24 Sep 2013 | United Kingdom | December 1938 | Darley Hall 182 Park St Lane, Park Street, St Albans, Hertfordshire, AL2 2AT |
| EMMETT, Bridget Anne, Professor RESIGNED | Director | 08 Sep 2021 | 10 Dec 2025 | Wales | December 1964 | Harpenden, Herts, AL5 2JQ |
| EVANS, David Anthony, Dr RESIGNED | Director | 30 Jun 2000 | 22 Nov 2006 | | November 1942 | Berry Hill, 6 Gong Hill Drive, Lower Bourne, Farnham, Surrey, GU10 3HG |
| FERGUSON, Iain George Thomas RESIGNED | Director | 08 Jul 1996 | 24 Feb 2005 | United Kingdom | July 1955 | Newlands Cottage 37 Trodds Lane, Merrow, Surrey, GU1 2XY |
| FLOWERS, Jonathan Philip Illtyd RESIGNED | Director | 18 Nov 2008 | 31 Dec 2012 | England | September 1964 | 2 Cowper Road, Harpenden, Hertfordshire, AL5 5NG |
| GODFRAY, Hugh Charles Jonathan, Professor RESIGNED | Director | 21 Aug 2013 | 31 Aug 2021 | England | October 1958 | Department Of Zoology, South Parks Road, Oxford, OX1 3PS, England |
| GRAHAM BRYCE, Ian James, Dr RESIGNED | Director | 30 Jun 2000 | 31 Jul 2004 | | March 1937 | 10 Watchbell Street, Rye, East Sussex, TN31 7HA |
| GRAY, Julie Elizabeth, Professor RESIGNED | Director | 10 Feb 2022 | 10 Jun 2026 | England | July 1962 | Harpenden, Herts, AL5 2JQ |
| GURR, Sarah Jane, Dr RESIGNED | Director | 30 Jun 2000 | 22 Nov 2006 | | May 1958 | 2 Stable Close, Finmere, Buckingham, Buckinghamshire, MK18 4AD |
| HAYWOOD, William John RESIGNED | Director | 04 Dec 2019 | 05 Dec 2023 | United Kingdom | January 1965 | Harpenden, Herts, AL5 2JQ |
| JAMES, Anthony Trafford RESIGNED | Director | | 24 May 1993 | | March 1922 | 9 High Street, Harrold, Bedford, Bedfordshire, MK43 7DQ |
| JARVIS, Stuart James RESIGNED | Director | 10 Feb 2015 | 07 Feb 2023 | England | November 1969 | Harpenden, Herts, AL5 2JQ |
| KILLHAM, Kenneth Stuart, Professor RESIGNED | Director | 30 Jun 2000 | 04 Mar 2009 | | March 1957 | Knowehead House Monymusk, Inverurie, Aberdeenshire, AB51 7SQ |
| LAWRENCE, David Kenneth, Dr RESIGNED | Director | 01 Jul 2007 | 12 May 2015 | United Kingdom | March 1949 | 135 Wilderness Road, Earley, Reading, Berkshire, RG6 5RD |
| LEAKE, Alastair Roland RESIGNED | Director | 12 May 2015 | 08 May 2019 | England | April 1961 | Harpenden, Herts, AL5 2JQ |
| LEGG, Brian James, Professor RESIGNED | Director | 30 Jun 2000 | 24 Nov 2008 | | July 1945 | 7 Blackmore Way, Blackmore End, Wheathampstead, Hertfordshire, AL4 8LU |
| LEONARD, Paul Keith RESIGNED | Director | 18 Nov 2008 | 05 Feb 2019 | United Kingdom | April 1962 | Edelweisslaan 53, Tervuren, 3080, Belgium |
| MANNING, Louise Joan, Professor RESIGNED | Director | 06 Dec 2019 | 09 Jul 2024 | United Kingdom | February 1964 | Harpenden, Herts, AL5 2JQ |
| MANNING, Louise Joan, Professor RESIGNED | Director | 04 Dec 2019 | 06 Dec 2019 | United Kingdom | February 1965 | Harpenden, Herts, AL5 2JQ |
| PAYNE, Christopher Charles, Professor RESIGNED | Director | 30 Jun 2000 | 03 Oct 2001 | | May 1946 | Old Thatch 15 North Street, Marcham, Abingdon, Oxfordshire, OX13 6NG |
| PEMBERTON, Antony Francis RESIGNED | Director | | 31 Jul 2004 | United Kingdom | February 1942 | Church Farm, Trumpington, Cambridge, Cambridgeshire, CB2 2LG |
| READ, David John RESIGNED | Director | 30 Jun 2000 | 01 Jun 2013 | United Kingdom | January 1939 | Minestone Cottage, Youlgrave, Bakewell, Derbyshire, DE45 1WD |
| RILEY, Ralph, Sir RESIGNED | Director | | 02 Jul 1999 | | October 1924 | 16 Gog Magog Way, Stapleford, Cambridge, Cambridgeshire, CB2 5BQ |
| ROBINSON, Carol Vivien, Professor RESIGNED | Director | 01 Oct 2004 | 09 May 2008 | | April 1956 | 7 Church Lane, Madingley, Cambridge, Cambridgeshire, CB3 8AF |
| SCRAGG, Neil Grant RESIGNED | Director | 10 May 2023 | 18 May 2026 | England | October 1968 | Harpenden, Herts, AL5 2JQ |
| SMITH, Sally Ann RESIGNED | Director | 01 Mar 2013 | 10 Feb 2021 | England | December 1973 | Harpenden, Herts, AL5 2JQ |
| TALBOT, Nicholas Jose, Professor RESIGNED | Director | 17 Jul 2007 | 13 May 2014 | England | September 1965 | 1 Eastdon Cottages, Starcross, Exeter, Devon, EX6 8RL |
| THOMAS, Sandra Mary, Professor RESIGNED | Director | 06 Sep 2022 | 25 Aug 2026 | England | June 1951 | Harpenden, Herts, AL5 2JQ |
| TURNER, Roger George, Dr RESIGNED | Director | 30 Jun 2000 | 24 Nov 2008 | Uk | September 1942 | 5 The Paddocks, Worlington, Bury St Edmunds, Suffolk, IP28 8SB |
| WINTER, David Michael, Professor RESIGNED | Director | 01 Sep 2014 | 05 Dec 2023 | England | November 1955 | Harpenden, Herts, AL5 2JQ |